The symbiosis between plants and complex communities of microbes is one of the most widespread and ecologically important host-microbiome associations. Plant microbiomes affect ecosystem functioning directly through nutrient cycling, and indirectly through their ability to influence host response to abiotic and biotic perturbations. Plant microbiomes have been found to be structured by both abiotic and biotic factors and thus susceptible to environmental changes. Recent studies have found evidence that environmental stress influences the plant microbiome which in some cases result in plant adaptation to stress. However, it is not clear if this is universal or specific to the stress, host plant, or ecosystem. Additionally, it is not known how specific microbial communities influence host acclimation and adaptation to changing environmental conditions. Here we will explore the complex relationships of microbes and plants and how environmental perturbations affect these relationships. We will examine various plant microbial interactions across plant species, plant developmental stages, and plant tissue types with diverse speakers at different stages in their careers. Talks will range from microbial community structure to nutrient cycling including warming experiments, CO₂ manipulations, and nutrient additions, all in the context of plant microbiomes in a changing environment.