This session will describe recent advances in the development of a systems biology of ecology based on applying network theory to the dynamics of ecosystem services including the provision of fish by fisheries, plant reproductive services by pollinators, and carbon sequestration by whole ecosystems. Each application area will highlight how profound ecological research challenges are met. For example, a fisheries application will integrate the evolution of fish towards smaller earlier maturing fish in the context of ecosystem management. Pollination applications will integrate the consumption of trophic resources and reproductive services as well as genomic elucidation of interactions. Overall, these presentations will show how mechanistic modeling of organisms and their interactions parameterized by powerful theory and extensive observations are advancing as has systems biology with networks at their conceptual core.